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In zoology, copulation is animal sexual behavior in which a male introduces sperm into the female's body, especially directly into her reproductive tract.[1][2] This is an aspect of mating. Many aquatic animals use external fertilization, whereas internal fertilization may have developed from a need to maintain gametes in a liquid medium in the Late Ordovician epoch.[citation needed] Internal fertilization with many vertebrates (such as all reptiles, some fish, and most birds) occurs via cloacal copulation, known as cloacal kiss (see also hemipenis), while most mammals copulate vaginally, and many basal vertebrates reproduce sexually with external fertilization.[3][4]