Lichen growth forms

The three main growth forms
Very intricately branched, greenish, bush-like lichen hanging from a thin twig
Fruticose (Usnea filipendula)

Lichens are symbiotic organisms made up of multiple species: a fungus, one or more photobionts (an alga and/or a cyanobacteria) and sometimes a yeast. They are regularly grouped by their external appearance – a characteristic known as their growth form. This form, which is based on the appearance of vegetative part of the lichen (its thallus), varies depending on the species and the environmental conditions it faces. Those who study lichens (lichenologists) have described a dozen of these forms: areolate, byssoid, calicioid, cladoniform, crustose, filamentous, foliose, fruticose, gelatinous, leprose, placoidioid and squamulose. Traditionally, crustose (flat), foliose (leafy) and fruticose (shrubby) are considered to be the three main forms. In addition to these more formalised, traditional growth types, there are a handful of informal types named for their resemblance to the lichens of specific genera. These include alectorioid, catapyrenioid, cetrarioid, hypogymnioid, parmelioid and usneoid.


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